MANUSCRIPTS
Carlen, E.J., Caizergues, A.E., Jagiello, Z., Kuzyo, H., Munshi-South, J., Alberti, M., Angeoletto, F., Bonilla-Bedoya, S., Booth, W., Charmantier, A., Cocciardi J.M., Cook, E.M., Gotanda, K.M., Govaert, L., Johnson, L.E., Li, D., Malesis, A.N., Martin, E., Marzluff, J.M., Mazurek, M., Miles, L.S., Phifer-Rixey, M., Salazar-Valenzuela, D., Savage, A., Snyder, R., Stadnicki, I., Vergeles, Y., Verrelli, B.C., Villar, D.A., Wilhelm-Solomon, M., Willmott, M., Winchell, K.M., Yates, S.S., Zhou, Y., Garroway, C.J., & Szulkin, M. (2025) Legacy effects of religion, politics and war on urban evolutionary biology. Nature Cities. html, pdf
McCluney, K.E., Deviche, P., Sweazea, K.L., Carlen, E.J., Clark (Brown), J.A.G., Grade, A., Haight, J.D., Niesner, C., Trego, S., & Weiss, K.C.B. (2025) An integrated social–ecological–evolutionary–phenotypic (SEEP) approach to understanding animal responses to urbanization. Biological Reviews. html, pdf
Lewis, D.E., Losos, J.B., & Carlen, E.J. (2025) Differential Influence of Environmental Factors on Pigeon Density and Distribution in European vs. North American City. Urban Naturalist. html, pdf
*Carlen, E.J., *Miles, L.S., Nassrullah, Munshi-South, J., Johnson, M.T.J. (2025) No detectable effect of urbanization on genetic drift or gene flow in specialist herbivorous insects of milkweed. PLoS ONE. html, pdf [*Co-first authors]
Kreling, S.E.S., Vance, S.E., & Carlen, E.J. (2025) Adaptation in the Alleyways: Candidate genes under potential selection in urban coyotes. Genome Biology and Evolution. html, pdf
*Carlen, E.J., *Estien, C.O., & Schell, C.J. (2024) Examining the influence of sociodemographics, residential segregation, and historical redlining on eBird and iNaturalist data disparities in three U.S. cities. Ecology and Society. html, pdf [*Co-first authors]
Savage, A., Willmott, M., Moreno, P., Li, D., Malesis A., Jagiełło, Z., Miles, L.S., Kuzyo, H. Román Palacios, C., Salazar, D., Verrelli, B., Winchell, K.M., Alberti, M., Carlen, E.J., Falvey, C., Johnson, L., Martin, E., Marzluff, J., Munshi-South, J., Phifer-Rixey, M., Stadnicki, I., Szulkin, M., Bonilla-Bedoya, S., Zhou, Y., Gotanda, K. (2024) Online toolkits for collaborative and inclusive global research in urban evolutionary ecology. Ecology and Evolution. html, pdf
Johnson, M.T.J., Arif, I., Marchetti, F., Munshi-South, J., Ness, R.W., Szulkin, M., Verrelli, B.C., Yauk, C.L., Anstett, D.N., Booth, W., Caizergues, A.E., Carlen, E.J., Dant, A., González, J., González Lagos, C., Oman, M., Phifer-Rixey, M., Rennison, D., Rosenberg, M.S., Winchell, K.M. (2024). Effects of urban-induced mutations on ecology, evolution and health. Nature Ecology & Evolution. html, pdf
*Williams, T.D., *Kreling, S.E.S., Stanton, L.A. Wilkinson, C.E., Estien, C.O., Schell, C.J., Carlen, E.J. (2024) Of Rarity and Symbolism: Understanding Human Perceptions of Charismatic Color Morphs. Human Ecology. html, pdf [*Co-first authors]
Caizergues, A.E., Santangelo, J.S., Ness, R.W., Angeoletto, F., Anstett, D.N., Anstett, J., Baena-Diaz, F., Carlen, E.J., Chaves, J.A., Comerford, M.S., Dyson, K., Falahati-Anbaran, M., Fellowes, M.D.E., Hood, G.R., Iñiguez-Armijos, C., Kooyers, N.J., Hodgins, K.A., Lázaro-Lobo, A., Moles, A., Munshi-South, J., Paule, J., Porth, I.M. , Santiago-Rosario, L.Y., Stack Whitney, K., Tack, A., Johnson, M.T.J. Does urbanization lead to parallel demographic shifts across the world in a cosmopolitan plant? Molecular Ecology. html, pdf
*Carlen, E.J., *Estien, C.O., Perkins, D., Goldstein, B.R., Caspi, T., Williams, T.D., Kreling, S.E.S., Hentati, Y., Stanton, L.A., Des Roches, S., Johnson, R., Young, A.N., Cooper, C.B., Schell, C.J. (2024). A framework for contextualizing social‐ecological biases in contributory science data. People and Nature. html, pdf [*Co-first authors]
Fusco, N.A., Cosentino, B.J., Gibbs, J.P., Allen, M.L., Blumenfeld, A.J., Boettner, G.H., Carlen, E.J., Collins, M., Dennison, C., DiGiacopo, D. and Drapeau Picard, A.P., (2024). Population genomic structure of a widespread, urban‐dwelling mammal: The eastern grey squirrel (Sciurus carolinensis). Molecular Ecology. html, pdf
Winchell, K., Aviles-Rodriguez, K., Carlen, E.J., Miles, L., Charmantier, A., De León, L., Gotanda, K., Rivkin, L.R., Szulkin, M. and Verrelli, B.. (2022) Moving past the challenges and misconceptions in urban adaptation research. Ecology & Evolution. html, pdf
Santangelo, J.S., Ness, R.W., Cohan, B., Fitzpatrick, C.R., Innes, S.G., Koch, S., Miles, L.S., Munim, S., Peres-Neto, P.R., Prashad, C., Tong, A.T., [et al., including Carlen, E.J.]. (2022) Global urbanization drives parallel environmental and evolutionary change. Science. html, pdf
Dunn, R.R., Burger, J.R., Carlen, E.J., Koltz, A.M., Light, J.E., Martin, R.A., Munshi-South, J., Nichols, L.M., Vargo, E.L., Yitbarek, S. and Zhao, Y., (2022) A Theory of City Biogeography and the Origin of Urban Species. Frontiers in Conservation Science. html, pdf
Carlen, E.J., Li, R., Winchell, K.M. (2021). Urbanization predicts flight initiation distance in feral pigeons (Columba livia) across New York City. Animal Behaviour. html, pdf
Fusco, N.A., Carlen, E.J., Munshi-South, J. (2021). Urban Landscape Genetics: Are Biologists Keeping Up with the Pace of Urbanization? Current Landscape Ecology Reports. html, pdf
Miles, L.S., Carlen, E.J., Winchell, K.M., & Johnson, M.T.J. (2020). Urban evolution comes into its own: Emerging themes and future directions of a burgeoning field. Evolutionary Applications. html, pdf
Puckett, E.E., Sherratt, E., Combs, M., Carlen, E.J., Harcourt‐Smith, W., & Munshi‐South, J. (2020) Variation in brown rat cranial shape shows directional selection over 120 years in New York City. Ecology and Evolution. html, pdf
Carlen, E.J. & Munshi-South, J. (2020) Widespread Genetic Connectivity of Feral Pigeons Across the Northeastern Megacity. Evolutionary Applications. html, pdf
Engel, L.D., Carlen, E.J., Losos, J.B., & Winchell, K.M. (2020). Eastern Gray Squirrels (Sciurus carolinensis) Differ in Abundance and Response to Humans Across Urban Habitats of St. Louis. Urban Naturalist. html, pdf
Winchell, K.M., Carlen, E.J., Puente‐Rolón, A.R., & Revell, L.J. (2018). Divergent habitat use of two urban lizard species. Ecology and Evolution. html, pdf
Carlen, E.J., Rathbun, G.B., Olson, L.E., Sabuni, C.A., Stanley, W.T., & Dumbacher, J.P. (2017). Reconstructing the molecular phylogeny of giant sengis (Macroscelidea; Macroscelididae; Rhynchocyon). Molecular Phylogenetics and Evolution. html, pdf
Dumbacher, J.P., Carlen, E.J., & G.B. Rathbun. (2016). Petrosaltator gen. nov., a new genus replacement for the North African sengi Elephantulus rozeti (Macroscelidea; Macroscelididae). Zootaxa. html, pdf
McCluney, K.E., Deviche, P., Sweazea, K.L., Carlen, E.J., Clark (Brown), J.A.G., Grade, A., Haight, J.D., Niesner, C., Trego, S., & Weiss, K.C.B. (2025) An integrated social–ecological–evolutionary–phenotypic (SEEP) approach to understanding animal responses to urbanization. Biological Reviews. html, pdf
Lewis, D.E., Losos, J.B., & Carlen, E.J. (2025) Differential Influence of Environmental Factors on Pigeon Density and Distribution in European vs. North American City. Urban Naturalist. html, pdf
*Carlen, E.J., *Miles, L.S., Nassrullah, Munshi-South, J., Johnson, M.T.J. (2025) No detectable effect of urbanization on genetic drift or gene flow in specialist herbivorous insects of milkweed. PLoS ONE. html, pdf [*Co-first authors]
Kreling, S.E.S., Vance, S.E., & Carlen, E.J. (2025) Adaptation in the Alleyways: Candidate genes under potential selection in urban coyotes. Genome Biology and Evolution. html, pdf
*Carlen, E.J., *Estien, C.O., & Schell, C.J. (2024) Examining the influence of sociodemographics, residential segregation, and historical redlining on eBird and iNaturalist data disparities in three U.S. cities. Ecology and Society. html, pdf [*Co-first authors]
Savage, A., Willmott, M., Moreno, P., Li, D., Malesis A., Jagiełło, Z., Miles, L.S., Kuzyo, H. Román Palacios, C., Salazar, D., Verrelli, B., Winchell, K.M., Alberti, M., Carlen, E.J., Falvey, C., Johnson, L., Martin, E., Marzluff, J., Munshi-South, J., Phifer-Rixey, M., Stadnicki, I., Szulkin, M., Bonilla-Bedoya, S., Zhou, Y., Gotanda, K. (2024) Online toolkits for collaborative and inclusive global research in urban evolutionary ecology. Ecology and Evolution. html, pdf
Johnson, M.T.J., Arif, I., Marchetti, F., Munshi-South, J., Ness, R.W., Szulkin, M., Verrelli, B.C., Yauk, C.L., Anstett, D.N., Booth, W., Caizergues, A.E., Carlen, E.J., Dant, A., González, J., González Lagos, C., Oman, M., Phifer-Rixey, M., Rennison, D., Rosenberg, M.S., Winchell, K.M. (2024). Effects of urban-induced mutations on ecology, evolution and health. Nature Ecology & Evolution. html, pdf
*Williams, T.D., *Kreling, S.E.S., Stanton, L.A. Wilkinson, C.E., Estien, C.O., Schell, C.J., Carlen, E.J. (2024) Of Rarity and Symbolism: Understanding Human Perceptions of Charismatic Color Morphs. Human Ecology. html, pdf [*Co-first authors]
Caizergues, A.E., Santangelo, J.S., Ness, R.W., Angeoletto, F., Anstett, D.N., Anstett, J., Baena-Diaz, F., Carlen, E.J., Chaves, J.A., Comerford, M.S., Dyson, K., Falahati-Anbaran, M., Fellowes, M.D.E., Hood, G.R., Iñiguez-Armijos, C., Kooyers, N.J., Hodgins, K.A., Lázaro-Lobo, A., Moles, A., Munshi-South, J., Paule, J., Porth, I.M. , Santiago-Rosario, L.Y., Stack Whitney, K., Tack, A., Johnson, M.T.J. Does urbanization lead to parallel demographic shifts across the world in a cosmopolitan plant? Molecular Ecology. html, pdf
*Carlen, E.J., *Estien, C.O., Perkins, D., Goldstein, B.R., Caspi, T., Williams, T.D., Kreling, S.E.S., Hentati, Y., Stanton, L.A., Des Roches, S., Johnson, R., Young, A.N., Cooper, C.B., Schell, C.J. (2024). A framework for contextualizing social‐ecological biases in contributory science data. People and Nature. html, pdf [*Co-first authors]
Fusco, N.A., Cosentino, B.J., Gibbs, J.P., Allen, M.L., Blumenfeld, A.J., Boettner, G.H., Carlen, E.J., Collins, M., Dennison, C., DiGiacopo, D. and Drapeau Picard, A.P., (2024). Population genomic structure of a widespread, urban‐dwelling mammal: The eastern grey squirrel (Sciurus carolinensis). Molecular Ecology. html, pdf
Winchell, K., Aviles-Rodriguez, K., Carlen, E.J., Miles, L., Charmantier, A., De León, L., Gotanda, K., Rivkin, L.R., Szulkin, M. and Verrelli, B.. (2022) Moving past the challenges and misconceptions in urban adaptation research. Ecology & Evolution. html, pdf
Santangelo, J.S., Ness, R.W., Cohan, B., Fitzpatrick, C.R., Innes, S.G., Koch, S., Miles, L.S., Munim, S., Peres-Neto, P.R., Prashad, C., Tong, A.T., [et al., including Carlen, E.J.]. (2022) Global urbanization drives parallel environmental and evolutionary change. Science. html, pdf
Dunn, R.R., Burger, J.R., Carlen, E.J., Koltz, A.M., Light, J.E., Martin, R.A., Munshi-South, J., Nichols, L.M., Vargo, E.L., Yitbarek, S. and Zhao, Y., (2022) A Theory of City Biogeography and the Origin of Urban Species. Frontiers in Conservation Science. html, pdf
Carlen, E.J., Li, R., Winchell, K.M. (2021). Urbanization predicts flight initiation distance in feral pigeons (Columba livia) across New York City. Animal Behaviour. html, pdf
Fusco, N.A., Carlen, E.J., Munshi-South, J. (2021). Urban Landscape Genetics: Are Biologists Keeping Up with the Pace of Urbanization? Current Landscape Ecology Reports. html, pdf
Miles, L.S., Carlen, E.J., Winchell, K.M., & Johnson, M.T.J. (2020). Urban evolution comes into its own: Emerging themes and future directions of a burgeoning field. Evolutionary Applications. html, pdf
Puckett, E.E., Sherratt, E., Combs, M., Carlen, E.J., Harcourt‐Smith, W., & Munshi‐South, J. (2020) Variation in brown rat cranial shape shows directional selection over 120 years in New York City. Ecology and Evolution. html, pdf
Carlen, E.J. & Munshi-South, J. (2020) Widespread Genetic Connectivity of Feral Pigeons Across the Northeastern Megacity. Evolutionary Applications. html, pdf
Engel, L.D., Carlen, E.J., Losos, J.B., & Winchell, K.M. (2020). Eastern Gray Squirrels (Sciurus carolinensis) Differ in Abundance and Response to Humans Across Urban Habitats of St. Louis. Urban Naturalist. html, pdf
Winchell, K.M., Carlen, E.J., Puente‐Rolón, A.R., & Revell, L.J. (2018). Divergent habitat use of two urban lizard species. Ecology and Evolution. html, pdf
Carlen, E.J., Rathbun, G.B., Olson, L.E., Sabuni, C.A., Stanley, W.T., & Dumbacher, J.P. (2017). Reconstructing the molecular phylogeny of giant sengis (Macroscelidea; Macroscelididae; Rhynchocyon). Molecular Phylogenetics and Evolution. html, pdf
Dumbacher, J.P., Carlen, E.J., & G.B. Rathbun. (2016). Petrosaltator gen. nov., a new genus replacement for the North African sengi Elephantulus rozeti (Macroscelidea; Macroscelididae). Zootaxa. html, pdf
POPULAR SCIENCE
Carlen, E.J. (2025). "War, politics and religion shape wildlife evolution in cities." The Conversation. (em Português)
Carlen, E.J. & Williams, T.D. (2024). "Animals that are all black or all white have reputations based on superstition − biases that have real effects." The Conversation.
Carlen, E.J. & Williams, T.D. (2024). "Animals that are all black or all white have reputations based on superstition − biases that have real effects." The Conversation.
If you are unable to access any of the above websites and would like a copy of my article(s) please contact me. I am happy to send you any article(s) at no cost.